Testing a keystone predator hypothesis.
  1. The sea star Pisaster ochraceous feeds preferentially on mussels but will consume other invertebrates.

  2. When Pisaster was removed from an intertidal zone, mussels eventually took over the rock face and eliminated most other invertebrates and algae. In a control area from which Pisaster was not removed, there was little change in species diversity.
In the absence of the sea star as a keystone predator, mussels become a dominant species in intertidal communities.

Another example of keystone species is the sea otter in kelp forest communities.